Plant Nutrition
Hort Diagrams/Pics
Water Analysis
Foliar Nutrition
Photosynthesis
PH
Hort Terms
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Plant Nutrition
1. 16 Essential Elements
2. Nutritional Deficiencies & Explanations
3. Definition of Parts-per-Million(PPM)
4. Cations in Water
| 1. 16 Essential Elements
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An element is not considered essential unless a deficiency of
it
makes it impossible for the plant to complete its life cycle; such deficiency
is specific to the element in question and can be prevented or corrected only
by
supplying this element; and the element is directly involved in the nutrition
of the
plant quite apart from possible effects in correcting some unfavorable
microbial or
chemical condition of the soil or other culture medium.
-D. I. ARNON
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There are at present 16 Plant Nutrients which are known to be essential for the
growth and reproduction of higher plants. These elements are: carbon,
hydrogen, oxygen, phosphorus, potassium, nitrogen, sulfur, calcium, iron,
magnesium, boron, manganese, copper, zinc, molybdenum, and chlorine. Other
nutrients which may be essential are colbalt, strontium, vanadium, silicon, and
nickel though these are not considered essential by the world community for all
plants and are not routinely applied as a fertilizer nutrient. As techniques
for evaluating the essentiality of trace elements improve, it is generally
believed that more elements will be added to the list of essential plant
nutrients.
Discoverer and Discoverer of Essentiality for the Essential
Elements (Glass, 1989).
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Element
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Discoverer
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Year
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Discoverer of Essentiality
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Year
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C
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**
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**
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De Saussure
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1804
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H
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Cavendish
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1766
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De Saussu re
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1804
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O
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Priestley
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1774
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De Saussure
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1804
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N
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Rutherford
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1772
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De Saussure
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1804
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P
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Brand
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1772
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Ville
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1860
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S
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**
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**
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vonSachs, Knop
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1865
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K
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Davy
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1807
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vonSachs, Knop
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1860
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Ca
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Davy
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1807
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vonSachs, Knop
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1860
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Mg
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Davy
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1808
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vonSachs, Knop
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1860
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Fe
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**
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**
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vonSachs, Knop
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1860
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Mn
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Scheele
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1774
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McHargue
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1922
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Cu
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**
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**
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Sommer
Lipman & MacKinnon
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1931
1931
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Zn
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**
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**
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Sommer & Lipman
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1926
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Mo
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Hzelm
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1782
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Amon & Stout
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1939
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B
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Gay Lussac & Thenard
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1808
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Sommer & Lipman
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1926
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Cl
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Scheel
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1774
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Stout
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1954
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2. Nutritional Deficiencies & Explanations
| Functions |
Deficiency symptoms |
Calcium (Ca)
1. Constituent of cell walls in the form of calcium pectate; necessary
for normal mitosis (cell division).
2. Helps in membrane stability, maintenance of chromosome structure.
3. Activator of enzymes (phospholipase, argine kinase, adenosine
triphosphates).
4. Acts as a detoxifying agent by neutralizing organic acids in plants.
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1. Calcium deficiencies are not often seen in the field because secondary
effects associated with high acidity limit growth.
2. The young leaves of new plants are affected first. These are
often distorted, small and abnormally dark green.
3. Leaves may be cup-shaped and crinkled and the terminal buds
deteriorate with some breakdown of petioles.
4. Root growth is markedly impaired; rotting of roots occurs.
5. Desiccation of growing points (terminal buds) of plants under severe
deficiency.
6. Buds and blossoms shed prematurely.
7. Stem structure weakened.
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Magnesium (Mg)
1. Constituent of chlorophyll molecule and therefore essential for
photosynthesis.
2. An activator of many enzyme systems involved in carbohydrate
metabolism, synthesis of nucleic acids, etc.
3. Promotes uptake and translocation of phosphorus.
4. Helps in movement of sugars within plant.
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1. Interveinal chlorosis, mainly of older leaves, producing a streaked or
patchy effect; with acute deficiency the affected tissue may dry up and die.
2. Leaves usually small, brittle in final stages and curve upwards at
margin.
3. In some vegetable plants, chlorotic spots between veins, and marbling
with tints of orange, red and purple.
4. Twigs weak and prone to fungus attack, usually premature leaf drop.
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Sulphur (S)
1. Constituent of sulphur-bearing amino acids.
2. Involved in the metabolic activities of vitamins, biotin, thiamine and
coenzyme A.
3. Aids stabilization of protein structure.
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1. Younger leaves turn uniformly yellowish green or chlorotic.
2. Shoot growth is restricted, flower production often indeterminate.
3. Stems are stiff, woody and small in diameter.
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Iron (Fe)
1. Necessary for the synthesis and maintenance of chlorophyll in plants.
2. Essential component of many enzymes.
3. Plays an essential role in nucleic acid metabolism affects RNA
metabolism or chloroplasts.
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1. Typical interveinal chlorosis; youngest leaves first affected, points
and margins of leaves keep their green color longest.
2. In severe cases, the entire leaf, veins and interveinal areas turn
yellow and may eventually become bleached.
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Manganese (Mn)
1. A catalyst in several enzymatic and physiological reactions in plants;
a constituent of pyruvate carboxylase.
2. Involved in the plant's respiratory process.
3. Activates enzymes concerned with the metabolism of nitrogen and
synthesis of chlorophyll.
4. Controls the redox potential in plant cells during the phases of light
and darkness.
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1. Chlorosis between the veins of young leaves, characterized by the
appearance of chlorotic and necrotic spots in the interveinal areas.
2. Greyish areas appear near the base of the younger leaves and become
yellowish to yellow orange.
3. Symptoms of deficiency popularly known in oats as "grey speck", in
field peas as "marsh spot", in sugarcane as "streak disease''.
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Boron (B)
1. Affects the activities of certain enzymes.
2. Ability to complex with various polyhydroxy-compounds.
3. Increases permeability in membrane and thereby facilitates
carbohydrate transport.
4. Involved in lignin synthesis and other reactions.
5. Essential for cell division.
6. Associated with the uptake of calcium and its utilization by plants.
7. Regulates potassium/calcium ration in plants.
8. Essential for protein synthesis.
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1. Death of growing plants (shoot tips).
2. The leaves have a thick texture, sometimes curling and becoming
brittle.
3. Flowers do not form and root growth is stunted.
4. "Brown heart" in root crops characterized by dark spots on the
thickest part of the root or splitting at center.
5. Fruits such as apples develop "internal and external cork" symptoms.
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Molybdenum (Mo)
1. Associated with nitrogen utilization and nitrogen fixation.
2. Constituent of nitrate reductase and nitrogenase.
3. Required by Rhizobia for nitrogen fixation.
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1. Chlorotic interveinal mottling of the lower leaves, followed by
marginal necrosis and infolding of the leaves.
2. In cauliflower, the leaf tissues wither leaving only the midrib and a
few small pieces of leaf blade ("whip-tail").
3. Molybdenum deficiency is markedly evident in leguminous plants.
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Chlorine (Cl)
1. A constituent of auxin chloroindole-3-acetic acid which in immature
seeds takes the place of indole acetic acid.
2. Constituent of many compounds found in fungi and bacteria.
3. Stimulates the activity of some enzymes and influences carbohydrate
metabolism and water holding capacity of plant tissue.
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1. Wilting of leaflet tips, chlorosis of leaves and finally bronzing and
drying. |
Nitrogen(N)
1. An important constituent of
chlorophyll,
protoplasm, protein and nucleic.
2. Increases growth and development of all living tissues.
3. Improves the quality of leafy vegetables and fodders and the protein
content of food grains.
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1. Stunted growth.
2. Appearance of a light-green to pale-yellow color on the older leaves,
starting from the tips. This is followed by death and/or dropping of the
older leaves depending upon the degree of deficiency.
3. In acute deficiency, flowering is greatly reduced.
4. Lower protein content.
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Phosphorus (P)
1. A constituent of phosphatides, nucleic acids, proteins, phospholipids
and coenzymes NAD, NADP and ATP.
2. Constituent of certain amino acids.
3. Necessary for cell division, a constituent of chromosomes; stimulates
root development.
4. Necessary for meristematic growth; seed and fruit development;
stimulates flowering.
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1. Overall stunted appearance, the mature leaves have characteristic dark
to blue-green coloration, restricted root development.
2. In acute deficiency, occasional purpling of leaves and stems; spindly
growth.
3. Delayed maturity and lack of or poor seed and fruit development.
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Potassium (K)
1. An activator of enzymes involved in photosynthesis and protein and
carbohydrate metabolism.
2. Assists carbohydrate translocation; synthesis of protein and
maintenance of its stability; membrane permeability and pH control; water
utilization by stomatal regulation.
3. Improves utilization of light during cool and cloudy weather and
thereby enhances plant ability to resist cold and other adverse
conditions.
4. Enhances the plant's ability to resist diseases.
5. Increases size of grains or seeds and improves the quality of fruits
and vegetables.
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1. Chlorosis along the leaf margins followed by scorching and browning of
tips of older leaves; these symptoms then gradually progress inwards.
2. Slow and stunted growth of plants.
3. Stalks weak, and plants lodge easily.
4. Shriveled seeds or fruits.
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Zinc(Zn)
1. Involved in the biosynthesis of indole acetic acid.
2. Essential component of a variety of metallo-enzymes-carbonic
anhydrase, alcohol dehydrogenase, etc.
3. Plays a role in nucleic acid and protein synthesis.
4. Assists the utilization of phosphorus and nitrogen in plants.
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1. Deficiency symptoms mostly appear on the 2nd or 3rd fully mature
leaves from the top of plants.
2. In maize, from light yellow striping to a broad band of white or
yellow tissue with reddish purple veins between the midrib and edges of the
leaf, occurring mainly in the lower half of the leaf.
3. In wheat, a longitudinal band of white or yellow leaf tissue, followed
by interveinal chlorotic mottling and white to brown necrotic lesions in the
middle of the leaf blade; eventual collapse of the affected leaves near the
middle.
4. In rice, after 15-20 days of transplanting, small scattered light
yellow spots appear on the older leaves which later enlarge, coalesce and turn
deep brown; the entire leaf becomes rust-brown in color and dries out within a
month.
5. In citrus, irregular interveinal chlorosis; terminal leaves become
small and narrowed (little-leaf); fruit-bud formation is severely reduced;
twigs die back.
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Copper(Cu)
1. Constituent of cytochrome oxidase and component of many enzymes -
ascorbic acid oxidase, phenolase, lactase, etc.
2. Promotes formation of vitamin A in plants.
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1. In cereals, yellowing and curling of the leaf blade, restricted ear
production and poor grain set, indeterminate tillering.
2. In citrus, die back of new growth; exanthema pockets of gum
develop between the bark and the wood; the fruit shows brown excretions.
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3. Definition of Parts Per Million
Parts Per Million
(ppm) -A notation for indicating small amounts of materials. The
expression gives the number of units by weight of the substance per million
weight units of another substance, such as oven-dry soil. The term may be
used to express the number of weight units of a substance per million weight
units of a solution. The approximate weight of soil is 2 million pounds
per acre-6 inches. Therefore, ppm X 2 equals pounds per acre-6 inches of
soil, or ppm X 4 equals pounds per acre-foot of soil.
Parts per million (ppm) - Unit of concentration used to
describe substance concentration in a million; may be expressed on a weight,
volume, or number basis. The metric equivalent is mg/l.
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PARTS PER MILLION
1 gram material applied 1,000,000 g
soil = 1 ppm
1,000 mg applied to 1,000,000 g soil = 1 ppm
| 1,000,000 g soil = 1,000,000 cc = 1,000,000 ml |
1 mg applied to 1,000 ml = 1 ppm
1 mg placed in 1,000 ml would equal one part per million
1 mg per 1,000 ml
= 1 ppm
1 mg per 1,000 cc = 1 ppm
1 mg per 1,000 g =
1 ppm
1) Units that are considered related in the following
manner:
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1 gram (g) = 1 cubic centimeter (cc) = 1 milliliter
(ml)
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| 6" azalea pot filled would contain approx. |
1,000 grams |
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1,000 ml |
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1,000 cc |
2) Parts per million = mg/liter
One acre furrow slice 6" deep weighs
approx. 2,000,000 pounds
If you apply 1 pound of material to one
acre, you would apply:
1/2 lb to 1,000,000 lbs soil
1/2 lb to 1,000,000 lbs soil
1 lb material to 2,000,000 lbs. soil = 2 ppm material
1 pound = 454.6 grams
454.6 (g) applied to one acre 2,000,000 lb soil would =
227.3 g material 1,000,000 lb soil
= 227.3 g material 1,000,000 lb soil
= 1/ 2 ppm material applied/acre.
3)
1 10
100 1,000
gal gal
gal gal
1 gal = 3.7854 Liters
ppm = mg/1000 ml
= mg/liter
As one liter = 1000 ml
1 gal = 3.7854 Liters
10 gal = 37.854 Liters
100 gal = 378.54 Liters
1000 gal = 3785.4 Liters
How can I make a solution of KNO3 that will contain 101 ppm of KNO3?
What is the formula weight of KNO3?
K = 39
N = 14
O x 3 = 16 x 3 = 48
101 is the
formula wt.
What is the definition of ppm?
ppm = mg/L
Take formula wt. of KNO3 in mg = 101 mg
Place in one liter and I would have
101 mg KNO3 in one liter would equal?
101 ppm KNO3
How many ppm K would I have in this one liter?
Formula wt. in mg of K in KNO3 = 39 mg
= 39 ppm of K
How many ppm of N in KNO3 in 1 Liter? N = 14 mg
= 14 ppm N in 101 mg KNO3/L
101 mg KNO3 = x mg
KNO3
Liter
10 liters
x = 1010
mg KNO3/10 liters
1 gal
= 10 gal
3.7854 L x L
x = 37.854 Liters
101 mg KNO3 = x mg KNO3
1L
37.854 L (10 gal)
x = 3823.254 mg KNO3/10 gal to get 101 ppm KNO3
39 mg K (in KNO3)
= x mg K (KNO3)
37.854 L (10 gal)
x = 1,476.306 mg K/10 gal
How do I get 100 ppm K/gal?
KNO3 Formula wt = 101 mg
= 39 mg K
39 mg K
= 100 mg K
101 mg KNO3 x
mg KNO3
x = 258.9 mg KNO3 to give 100 ppm
K in 1 Liter
258.9 mg KNO3
= x mg KNO3
L
3.7854 L
x =
980 mg KNO 3 to get 100 ppm K in one gal
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4. Cations in Water
Table 17 (Major Cations In Water)
Constituents in Irrigation Water ---Major Cations
| Constituent |
Symbol |
Equivalent
Weight |
Desirable
Range (ppm) |
Desirable
Range (meq/L) |
Comments |
| Calcium |
Ca++ |
20.04 |
40-120 |
0.2-6.0 |
Essential plant nutrient, occurs naturally in most waters. Low levels
increase potential for Calcium deficient plants, but high levels are not
normally harmful. |
| Magnesium |
Mg++ |
12.15 |
6-24 |
0.5-2.0 |
Essential plant nutrient, occurs naturally in most waters. Low levels
increase potential for Magnesium deficient plants, but relatively high levels
are not normally a problem. |
| Sodium |
Na+ |
23.00 |
<50 |
<20 |
A non-essential nutrient, occurs naturally in most waters. Can influence
soil structure and plant uptake of Ca and Mg (see Sodium Adsorption Ratio
below). |
| Potassium |
K+ |
39.10 |
trace |
trace |
An essential nutrient, normally found in waters in small amounts.
Presence of more than trace amounts indicates waters may contain fertilizer. |
| Ammonium |
NH4+ |
17.03 |
trace |
trace |
A source of Nitrogen (an essential plant nutrient). Normally present in
waters in small amounts. More than trace amounts indicates water may
contain fertilizer. |
Table 18 (Major Anions In Ground Water)
Constituents in Irrigation Water ---Major Anions
| Constituent |
Symbol |
Equivalent Weight |
Desirable
Range (ppm) |
Desirable
Range (meq/L) |
Comments |
| Sulfate |
SO4- |
48.0 |
25-240 |
0.5-5.0 |
Contains Sulfate, an essential nutrient, occurs naturally in most waters.
Low levels increase potential for sulfur deficiency, but high levels are not
normally harmful. |
| Chloride |
C1- |
35.46 |
<70 |
<2.0 |
Occurs normally in most waters. Plants require trace amounts. High
concentrations are toxic to sensitive plants. |
| Phosphate |
PO4- |
94.97 |
trace |
trace |
Contains phosphorus an essential nutrient, normally found in waters only in
trace amounts. The presence of more than small amounts indicates water
may contain fertilizer or detergent. |
| Nitrate |
NO3- |
62.01 |
<10 |
0.12 |
Source of Nitrogen, an essential nutrient, occurs normally only in small
amounts. Presence of more than trace amounts indicates water may contain
fertilizer. The EPA Drinking Water Standard is 10 ppm maximum. |
Table 19 (Boron And Flouride)
Boron and Fluoride
| Constituents |
Symbol |
Desired Range (ppm) |
Comments |
| Boron |
B |
<0.5 |
An essential plant nutrient, occurs normally in most water. Low levels
increase potential for Boron Deficiency. High levels can be toxic to
plants. When evaluating Boron fertility, remember to include Boron
supplied by fertilizer and growing medium. |
| Fluoride |
F |
<0.75 |
A non-essential nutrient that occurs in some waters and is often added to
public drinking supplies. High Fluoride concentrations can cause toxicity
in sensitive plants. |
Table 20 (Irrigation Water Classifications)
Qualitative Classification of Irrigation Waters
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Class 1 -
Excellent to Good |
Class 2 -
Good to Injurious |
Class 3 -
Injurious to Unsatisfactory |
| EC, ds / m |
Less than 1.0 |
1.0 - 3.0 |
More than 3.0 |
| Boron, ppm |
Less than 0.5 |
0.5 - 2.0 |
More than 2.0 |
| Sodium, percent |
Less than 60 |
60 - 75 |
More than 75 |
| Chloride, me/L |
Less than 5 |
5 - 10 |
More than 10 |
Table 21 (Water Quality Indices)
Maximum Acceptable Water Quality - Indices for Bedding Plants
| Variable |
Plug Production |
Finish Flats and Pots |
| pH1 (acceptable range) |
5.5 to 7.5 |
5.5 to 7.5 |
| Alkalinity2 |
1.5 me/l |
2.0 me/l |
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(75 ppm) |
(100 ppm) |
| Hardiness3 |
3.0 me/l |
3.0 me/l |
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(150 ppm) |
(150 ppm) |
| EC |
1.0 mS |
1.2 mS |
| Ammonium-N |
20 ppm |
40 ppm |
| Boron |
0.5 ppm |
0.5 ppm |
Adapted from P.V. Nelson, Fertilization. Pp. 151-176. In:E. J. Holcomb
(ed). Bedding Plants IV. A Manual on the Culture of Bedding Plants as a
Greenhouse Crop, Ball Publishing, Batavia, IL (1994). Used with permission.
1pH not very important alone; alkalinity level more important
2Moderately higher alkalinity levels are acceptable
when lower amounts of limestone are incorporated into the substrate during
its formulation. Very high alkalinity levels require acid injection into
water source.
3High hardness values are not a problem if calcium and magnesium
concentrations are adequate and soluble salt level is tolerable.
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